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Either the festival is a financial success, or the parks Department will be eliminated.
After their bad date, Ann apologizes to Chris and asks to go out again, and he accepts.
Meanwhile, Andy gets help from Chris in winning back April, which makes Ann worry that Andy will once again interfere in her relationship.
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Leslie (Amy Poehler) and the parks department return to work after three months, only to be told by state auditors Chris (Rob Lowe) and Ben (Adam Scott) that the budget crisis still looms.
After unsuccessfully trying to have their budget restored by using Ann (Rashida Jones) to seduce Chris with a date, Leslie decides to make an all or nothing play by resurrecting the Pawnee harvest festival.
Eventually Leslie holds a public forum to decide what will go in the capsule, but it grows quickly out of control when the Pawnee citizens cannot agree on what it should include.
Leslie resolves the matter by including nothing but a video-recording of the meeting, which she feels captures Pawnee by showing the passion of its citizens.
Instead, Ron and Tammy reconnect and end up remarrying each other in the middle of a night of drinking and wild sex that leaves them both in prison.
Leslie asks Pawnee police Chief Hugh Trumple (Eric Pierpoint) to release Ron as a favor, even though the favor she originally planned to ask was for the police to volunteer officers for security as the harvest festival.
April, still angry with Andy for having kissed Ann, returns from a vacation with a new boyfriend from Venezuela, Eduardo. Flu season strikes Pawnee, and Ann must care for the flu victims at the hospital, which include April, Leslie, and Chris.
While April tries to make life hard for Ann as revenge for kissing Andy, Ron needs an assistant to fill in for April and selects Andy, whom he bonds with over the Colts, lunch, and playing football in the parking lot.
Just as Ron and Wendy break up when Wendy decides to move back to Canada, Ron's ex-wife Tammy (Megan Mullally) returns to try to make Ron's life miserable.